Emma Sing comes successful astatine full-back successful Kildunne's spot for Saturday's rematch (00:00 BST) against Canada successful Ottawa, portion Tatyana Heard and Claudia Moloney-MacDonald - backline starters successful the decision - driblet retired of the matchday squad entirely.
Helena Rowland and Mia Venner regenerate Heard and Moloney-MacDonald astatine wrong centre and helping respectively.
In the 4th alteration to the starting line-up, Sadia Kabeya, who appeared disconnected the seat past week connected her comeback from a limb injury, starts with Alex Matthews, two-time Rugby World Cup winner, dropping to the bench.
Exeter's Flo Robinson and Sale's Hollie Aitchison are included connected the seat arsenic a new-look acceptable of half-back replacements.
Sing, a rangy runner who tin besides footwear goals, started 2 matches astatine full-back during England's Six Nations campaign, with Kildunne switching to a antithetic presumption connected the helping connected each occasion.
Sing's Gloucester-Hartpury team-mate Venner made her England debut successful 2020 aged conscionable 17.
Venner has won lone different 4 caps since but was putting unit connected England's much established backmost 3 astatine the commencement of this year, appearing successful wins implicit Scotland and Italy.
The 24-year-old finished arsenic the PWR's associated apical try-scorer past season, crossing 12 times.
Moloney-MacDonald outgo her broadside an aboriginal effort successful the decision by Canada when, aft doing superbly to prime Jess Breach's walk disconnected her toes, she spilled the shot implicit the enactment successful the sixth minute.
"We've earned a beardown play but thing astir it is owed to america going forward," said Mitchell.
"Friday [Saturday 00:00 BST] is an accidental to use sustained pressure, larn from what we missed and travel retired the different broadside amended than we went in."
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